"other (evolutionary) branch"  so-named because it represents a distinct branch of tyrannosaur evolution, retaining some primitive features (a long snout and more teeth). The maxila has 16, possibly 17 teeth, and the dentary has 18 teeth, compared with a maximum of 13 maxillary and 15 dentary teeth in Tarbosaurus. The type specimen has a beautifully preserved braincase with abroad nuchal crest and a downturned occiput.